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(!B3 I{5upisoelectric point, )Q:NpI 0"$f8 2ĉ[aaQ5uw:N:NevnmpHv:NaavI{5up.xfL *#5 I{5upv{&  $ 1u_Sw㉻y8^pe*    L 1udkSw ]w(WpIe [R+]= [R-] 4" tetS_'0  $NS[pe -lg [H+ ] 2 = - lg K1 - lg K2 5" -lg [H+ ] = pH - lg K1 = pK1 - lg K2 = pK2 4" 2 pH = pK1 + pK2 N pI:NI{5upevpH R pI =  pK1 +pK2 GlypI = 2 .4 + 9.6) = 5.97N                        #V1udkSw: I{5upvS_Naa$N'`yP[r`$NO WVpKevCO2S(uKmSlKmϑ {QSRS^vaaϑ0 c0ˆ}v(TYNSS^ FOupOe^]0Pr; 2 -& vQNSf[S^$ N2uS^ NN]xxvS^ NN]xWlDNFB vS^ _kx0lxovS^ C-zKm[ P56-57 NgIQvS^ N5 5 -SkxW-S(2-]xW2ux)vS^UU@ NN]xxvS^$XHN2 CH COOH +HNO2 R-CH-COOH + N2 R CH +H2O p 2    K *Nё^\yP[S^ ~B\g  2$V-Nv(lWx]^[teRP[ y(lWxkW(animo acid residue);6,$)Y6qX[(Wv;m'`$ (6 ;m'`(active peptide)X[(WNuirSOQ wQTyyrkuirf[Rv0 (O(;(;71mQ0ˆ}v(v~g" "( gWc(WzSO_gSO-NSNSP[bWV(WzzvST0 gacُNSNWV(WUS.eleSb_bv N TvzSO_gSO0 ˆ}v(vgacˆ}v(yr gvzz~g09n  M83N ˆ}v(vN~~g" $ $  ˆ}v(vN~~g/fN.ޏcaav~'`^R0 10T.v~g peptide)N*N(lWxv-WTSN(lWxv-(lW14l)T bvSTir0h'Z,Zff  !l 20vW,gޏce_  .( .(peptide bond):N1*N(lWxv-(lWTSN(lWxv-WKN14lTb_bvqQN. sSp.0 dk:Nˆ}v(-N(lWxޏcvW,ge_0(KZ;T  &z{ 1 .wQؚ^3z['` 2 wQS.'`( N1ulR 3 USCQHTs^bSTR'`'`( 4 C=ON NHS_cR0 AA6- yz 30/Qvޏc  Nkx.  i_1u-Nv^MO N$N*NJS(l x1"lޏc b dky(lxk W /fޏcQbv;N eh.0 R3z[zz~gKNN[u ir;mR0 :f ?:"Nkx.vޏc y{| gYy S'`BR$  @; O\ }"$0 -N\-~ހRlvNyˆ}v(o }0 RMNO@|0 ~g A21aaB30aa g$N[Nkx. (WAQaa6Taa11 NNkx.ޏb\s0Y[r\ } txH '@Ypolypeptide)Y*N(lWxN.ޏce_b_b. 8!$$$ fNQ`N`]NSCNz/f8ny(lWSNz:N8nyW0 -Nv(lWx]^[teRP[ y(lWxkW(animo acid residue);^HZ!F&C=N ˆ}v(vzz~g  $ li_Prozz~gSyProvgaؚ~~g cProRP[-N@b gSP[TWV(W N~zzvcRSvpT0.7H 1 ~~rˆ}vRP[(  $ Xy{|:҉ˆ}v0Sˆ}v0@~~ˆ}v0~~ˆ}v0 R:(WuirSOQOb0/ec0~R0>-Zff -2 tˆ}vRP[: $($ ^y{|@~ˆ}v0~ˆ}v0nˆ}v0t ˆ}v0vˆ}v0o }ˆ}v0bSOI{020 f-$ 0DA10ˆ}v(RP[-Nv^qQN.$ 1 "l. W Nv'lNN(lW Nv"lSP[@bb_b0 \O(uQ~cN~~g:N N0 V~~g@b0 2 u4l.aa^g'`Ob_b0 \O(u~c N~~g0RZT &\EB 3 v.:$$  P&^ck5uwvOǏY5u_Rb_b0 4 _NSR -N'`RP[bSP[v\O(uR0J   )!k~.vyrpN\O(u $ >.1_ peϑY0 @bN (W~cˆ}v(zz~g-Nw͑\O(u0   NL20ˆ}v(vN~~g  $ ^cˆ}v(Y,gvbSTv~b__ - e~g - bS~g - l҉~g 1uVl$0@ OO&1 - e(-helix) "$$, nV-NaakW-Nv-NHNMR N*NaavC=OSb_b"l. O_HTervf0 "l.STQNNN-N_ts^L0P8$d!    PM42 -bS(-pleated sheet) ,$$$4`~gyrp 8OU\ cB\cR, (Wv N T:Skb N Tb_b"l.,~c~gv3z['`0<1f   1XX-bS g$Ny{|W  $`s^L@b gvNzYN TNz "l. NNpTWv0Y - ҉ˆ}v0 1Z16TP"3 - l҉(- turn)$, bQb_b"l. 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N0ˆ}v(vuirf[R"  "66 H10RY7hˆ}v(RvY7h'` 1)0PS\O(u QNN@b gvv/fˆ}v(&Z Z%$ % 6 0o }\O(u \ }I{ 7 0cST OOo`vSSO SSOˆ}vI{ 8 0c6R~ހu0RS uVP[0;Oˆ}v 9 0%{Q\O(u0_;mRI{ebw͑\O(u0 20R YBgˆ}v(Rv YBg'` vPSRSYV }vq_T v^hsQzN'`0S'`I{ " dZ $ $ $  $($$)bN0ˆ}v(vR{|  "(  R{|SR cRP[b_r0RP[~b0n㉦^0uirR N cRP[b_rtrˆ}vT~~rˆ}v N cRP[~b{USˆ}vT~Tˆ}v  N cn㉦^nˆ}v0|ˆ}v0~ˆ}v0tˆ}v07ˆ}v0nˆ}v0lxˆ}vI{0 kZf f f k(V) cuirR* f;m'`ˆ}v(:Sb(Wu}TǏ z-NNR g;m'`vˆ}v(SvQMRSO0Y:o }ˆ}v(0Џˆ}v(0Obb2_ˆ}v(0kˆ}v(I{0 ^;m'`ˆ}v(ُ{|Wˆ}v([uirSOwObb/ec\O(u0Ylxˆ}v0҉ˆ}v0`Z$  $$  _  N0ˆ}v(v~b  "( N ˆ}v(vCQ }~b C0H0O0N S\ϑv S yrpN (WTyˆ}v(-Nvs^GW+Tϑ:N16% (u[.llSKm[ˆ}v(+Tϑ ˆ}v(+Tϑ(g) = ˆ}v(+T.lϑ6.25ff*$b@  V0(lWxvSf[  "( z(lWxamino acid, {Qaa) ˆ}v(vW,g~bUSMO /f+T(lWvx0 )Y6qˆ}v(1u20y(lWx~b0X$$9$$8 $N (lWxv~g  (  10_((( 20 -(lWxvgW  $gW1uNWVbSP[(WzzcRv N [y'` _wvIQf[;m'`zSO~g yzSOgW0 zSOgW (lW(W]:NL-W (lW(WS:ND-W/f=$f.N  6)Y6qpr-NvaaGW:NL-W $NWaavutR N T0 2^ v30 - (lWxwQeIQ'`du(lxY J< ( ((- $6  & gWNkeIQ^evc[^sQ|( N (lWxvy{|S~g"    010 c~gSR:N 4'Y{| 1 eO 6 $   )YQ(lx 7(lx )YQp 7(lp BBZ>( B2 eO ( t N(lx j(lx &; : ;3 Bgs(lWx   4 BgsN(lWx    #20 cRWVvg'`R"    g'`(lWx g'` N&^5uwN0ς0)YQp0 7(lp0j0JS0 g'`&^ck5uw)YQ070 g'`&^5uw~0V0|0g3 g$30 cNSO/f&TSTbegR  _(lWx:gSO NꁫTb _{1uߘirO~v(lWx0 V0r0ς0,02ukx0N0_N0N JS_(lWxNSOSTb FO|^?QgTb^ Nn0 ~*0|*0 ^_(lWx:gSOSꁫTbv(lWx0   q40^ˆ}v((lWxB &( $^ˆ}v((lWx_NwQ guir;m'`0P456$& N (lWxv͑tS'`(& $ 10N,irt'`( 20$N'`㉻yNI{5up$N'`'`(  1 (lWx:N$N'`5u( RM  # M' (2 aa㉻ye_SQN@bYsXvpH&$.  (!B3 I{5upisoelectric point, )Q:NpI 0"$f8 2ĉ[aaQ5uw:N:NevnmpHv:NaavI{5up.xfL *#5 I{5upv{&  $ 1u_Sw㉻y8^pe*    L 1udkSw ]w(WpIe [R+]= [R-] 4" tetS_'0  $NS[pe -lg [H+ ] 2 = - lg K1 - lg K2 5" -lg [H+ ] = pH - lg K1 = pK1 - lg K2 = pK2 4" 2 pH = pK1 + pK2 N pI:NI{5upevpH R pI =  pK1 +pK2 GlypI = 2 .4 + 9.6) = 5.97N                        #V1udkSw: I{5upvS_Naa$N'`yP[r`$NO WVpKevCO2S(uKmSlKmϑ {QSRS^vaaϑ0 c0ˆ}v(TYNSS^ FOupOe^]0Pr; 2 -& vQNSf[S^$ N2uS^ NN]xxvS^ NN]xWlDNFB vS^ _kx0lxovS^ C-zKm[ P56-57 NgIQvS^ N5 5 -SkxW-S(2-]xW2ux)vS^UU@ NN]xxvS^$XHN2 CH COOH +HNO2 R-CH-COOH + N2 R CH +H2O p 2    K *Nё^\yP[S^ ~B\g  2$V-Nv(lWx]^[teRP[ y(lWxkW(animo acid residue);6,$)Y6qX[(Wv;m'`$ (6 ;m'`(active peptide)X[(WNuirSOQ wQTyyrkuirf[Rv0 (O(;(;71mQ0ˆ}v(v~g" "( gWc(WzSO_gSO-NSNSP[bWV(WzzvST0 gacُNSNWV(WUS.eleSb_bv N TvzSO_gSO0 ˆ}v(vgacˆ}v(yr gvzz~g09n  M83N ˆ}v(vN~~g" $ $  ˆ}v(vN~~g/fN.ޏcaav~'`^R0 10T.v~g peptide)N*N(lWxv-WTSN(lWxv-(lW14l)T bvSTir0h'Z,Zff  !l 20vW,gޏce_  .( .(peptide bond):N1*N(lWxv-(lWTSN(lWxv-WKN14lTb_bvqQN. sSp.0 dk:Nˆ}v(-N(lWxޏcvW,ge_0(KZ;T  &z{ 1 .wQؚ^3z['` 2 wQS.'`( N1ulR 3 USCQHTs^bSTR'`'`( 4 C=ON NHS_cR0 AA6- yz 30/Qvޏc  Nkx.  i_1u-Nv^MO N$N*NJS(l x1"lޏc b dky(lxk W /fޏcQbv;N eh.0 R3z[zz~gKNN[u ir;mR0 :f ?:"Nkx.vޏc y{| gYy S'`BR$  @; O\ }"$0 -N\-~ހRlvNyˆ}v(o }0 RMNO@|0 ~g A21aaB30aa g$N[Nkx. 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N0ˆ}v(vuirf[R"  "66 H10RY7hˆ}v(RvY7h'` 1)0PS\O(u QNN@b gvv/fˆ}v(&Z Z%$ % 6 0o }\O(u \ }I{ 7 0cST OOo`vSSO SSOˆ}vI{ 8 0c6R~ހu0RS uVP[0;Oˆ}v 9 0%{Q\O(u0_;mRI{ebw͑\O(u0 20R YBgˆ}v(Rv YBg'` vPSRSYV }vq_T v^hsQzN'`0S'`I{ " dZ $ $ $  $($$)bN0ˆ}v(vR{|  "(  R{|SR cRP[b_r0RP[~b0n㉦^0uirR N cRP[b_rtrˆ}vT~~rˆ}v N cRP[~b{USˆ}vT~Tˆ}v  N cn㉦^nˆ}v0|ˆ}v0~ˆ}v0tˆ}v07ˆ}v0nˆ}v0lxˆ}vI{0 kZf f f k(V) cuirR* f;m'`ˆ}v(:Sb(Wu}TǏ z-NNR g;m'`vˆ}v(SvQMRSO0Y:o }ˆ}v(0Џˆ}v(0Obb2_ˆ}v(0kˆ}v(I{0 ^;m'`ˆ}v(ُ{|Wˆ}v([uirSOwObb/ec\O(u0Ylxˆ}v0҉ˆ}v0`Z$  $$  _  N0ˆ}v(v~b  "( N ˆ}v(vCQ }~b C0H0O0N S\ϑv S yrpN (WTyˆ}v(-Nvs^GW+Tϑ:N16% (u[.llSKm[ˆ}v(+Tϑ ˆ}v(+Tϑ(g) = ˆ}v(+T.lϑ6.25ff*$b@  V0(lWxvSf[  "( z(lWxamino acid, {Qaa) ˆ}v(vW,g~bUSMO /f+T(lWvx0 )Y6qˆ}v(1u20y(lWx~b0X$$9$$8 $N (lWxv~g  (  10_((( 20 -(lWxvgW  $gW1uNWVbSP[(WzzcRv N [y'` _wvIQf[;m'`zSO~g yzSOgW0 zSOgW (lW(W]:NL-W (lW(WS:ND-W/f=$f.N  6)Y6qpr-NvaaGW:NL-W $NWaavutR N T0 2^ v30 - (lWxwQeIQ'`du(lxY J< ( ((- $6  & gWNkeIQ^evc[^sQ|( N (lWxvy{|S~g"    010 c~gSR:N 4'Y{| 1 eO 6 $   )YQ(lx 7(lx )YQp 7(lp BBZ>( B2 eO ( t N(lx j(lx &; : ;3 Bgs(lWx   4 BgsN(lWx    #20 cRWVvg'`R"    g'`(lWx g'` N&^5uwN0ς0)YQp0 7(lp0j0JS0 g'`&^ck5uw)YQ070 g'`&^5uw~0V0|0g3 g$30 cNSO/f&TSTbegR  _(lWx:gSO NꁫTb _{1uߘirO~v(lWx0 V0r0ς0,02ukx0N0_N0N JS_(lWxNSOSTb FO|^?QgTb^ Nn0 ~*0|*0 ^_(lWx:gSOSꁫTbv(lWx0   q40^ˆ}v((lWxB &( $^ˆ}v((lWx_NwQ guir;m'`0P456$& N (lWxv͑tS'`(& $ 10N,irt'`( 20$N'`㉻yNI{5up$N'`'`(  1 (lWx:N$N'`5u( RM  # M' (2 aa㉻ye_SQN@bYsXvpH&$.  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N0ˆ}v(vuirf[R"  "66 H10RY7hˆ}v(RvY7h'` 1)0PS\O(u QNN@b gvv/fˆ}v(&Z Z%$ % 6 0o }\O(u \ }I{ 7 0cST OOo`vSSO SSOˆ}vI{ 8 0c6R~ހu0RS uVP[0;Oˆ}v 9 0%{Q\O(u0_;mRI{ebw͑\O(u0 20R YBgˆ}v(Rv YBg'` vPSRSYV }vq_T v^hsQzN'`0S'`I{ " dZ $ $ $  $($$)bN0ˆ}v(vR{|  "(  R{|SR cRP[b_r0RP[~b0n㉦^0uirR N cRP[b_rtrˆ}vT~~rˆ}v N cRP[~b{USˆ}vT~Tˆ}v  N cn㉦^nˆ}v0|ˆ}v0~ˆ}v0tˆ}v07ˆ}v0nˆ}v0lxˆ}vI{0 kZf f f k(V) cuirR* f;m'`ˆ}v(:Sb(Wu}TǏ z-NNR g;m'`vˆ}v(SvQMRSO0Y:o }ˆ}v(0Џˆ}v(0Obb2_ˆ}v(0kˆ}v(I{0 ^;m'`ˆ}v(ُ{|Wˆ}v([uirSOwObb/ec\O(u0Ylxˆ}v0҉ˆ}v0`Z$  $$  _  N0ˆ}v(v~b  "( N ˆ}v(vCQ }~b C0H0O0N S\ϑv S yrpN (WTyˆ}v(-Nvs^GW+Tϑ:N16% (u[.llSKm[ˆ}v(+Tϑ ˆ}v(+Tϑ(g) = ˆ}v(+T.lϑ6.25ff*$b@  V0(lWxvSf[  "( z(lWxamino acid, {Qaa) ˆ}v(vW,g~bUSMO /f+T(lWvx0 )Y6qˆ}v(1u20y(lWx~b0X$$9$$8 $N (lWxv~g  (  10_((( 20 -(lWxvgW  $gW1uNWVbSP[(WzzcRv N [y'` _wvIQf[;m'`zSO~g yzSOgW0 zSOgW (lW(W]:NL-W (lW(WS:ND-W/f=$f.N  6)Y6qpr-NvaaGW:NL-W $NWaavutR N T0 2^ v30 - (lWxwQeIQ'`du(lxY J< ( ((- $6  & gWNkeIQ^evc[^sQ|( N (lWxvy{|S~g"    010 c~gSR:N 4'Y{| 1 eO 6 $   )YQ(lx 7(lx )YQp 7(lp BBZ>( B2 eO ( t N(lx j(lx &; : ;3 Bgs(lWx   4 BgsN(lWx    #20 cRWVvg'`R"    g'`(lWx g'` N&^5uwN0ς0)YQp0 7(lp0j0JS0 g'`&^ck5uw)YQ070 g'`&^5uw~0V0|0g3 g$30 cNSO/f&TSTbegR  _(lWx:gSO NꁫTb _{1uߘirO~v(lWx0 V0r0ς0,02ukx0N0_N0N JS_(lWxNSOSTb FO|^?QgTb^ Nn0 ~*0|*0 ^_(lWx:gSOSꁫTbv(lWx0   q40^ˆ}v((lWxB &( $^ˆ}v((lWx_NwQ guir;m'`0P456$& N (lWxv͑tS'`(& $ 10N,irt'`( 20$N'`㉻yNI{5up$N'`'`(  1 (lWx:N$N'`5u( RM  # M' (2 aa㉻ye_SQN@bYsXvpH&$.  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N0ˆ}v(vuirf[R"  "66 H10RY7hˆ}v(RvY7h'` 1)0PS\O(u QNN@b gvv/fˆ}v(&Z Z%$ % 6 0o }\O(u \ }I{ 7 0cST OOo`vSSO SSOˆ}vI{ 8 0c6R~ހu0RS uVP[0;Oˆ}v 9 0%{Q\O(u0_;mRI{ebw͑\O(u0 20R YBgˆ}v(Rv YBg'` vPSRSYV }vq_T v^hsQzN'`0S'`I{ " dZ $ $ $  $($$)bN0ˆ}v(vR{|  "(  R{|SR cRP[b_r0RP[~b0n㉦^0uirR N cRP[b_rtrˆ}vT~~rˆ}v N cRP[~b{USˆ}vT~Tˆ}v  N cn㉦^nˆ}v0|ˆ}v0~ˆ}v0tˆ}v07ˆ}v0nˆ}v0lxˆ}vI{0 kZf f f k(V) cuirR* f;m'`ˆ}v(:Sb(Wu}TǏ z-NNR g;m'`vˆ}v(SvQMRSO0Y:o }ˆ}v(0Џˆ}v(0Obb2_ˆ}v(0kˆ}v(I{0 ^;m'`ˆ}v(ُ{|Wˆ}v([uirSOwObb/ec\O(u0Ylxˆ}v0҉ˆ}v0`Z$  $$  _  N0ˆ}v(v~b  "( N ˆ}v(vCQ }~b C0H0O0N S\ϑv S yrpN (WTyˆ}v(-Nvs^GW+Tϑ:N16% (u[.llSKm[ˆ}v(+Tϑ ˆ}v(+Tϑ(g) = ˆ}v(+T.lϑ6.25ff*$b@  V0(lWxvSf[  "( z(lWxamino acid, {Qaa) ˆ}v(vW,g~bUSMO /f+T(lWvx0 )Y6qˆ}v(1u20y(lWx~b0X$$9$$8 $N (lWxv~g  (  10_((( 20 -(lWxvgW  $gW1uNWVbSP[(WzzcRv N [y'` _wvIQf[;m'`zSO~g yzSOgW0 zSOgW (lW(W]:NL-W (lW(WS:ND-W/f=$f.N  6)Y6qpr-NvaaGW:NL-W $NWaavutR N T0 2^ v30 - (lWxwQeIQ'`du(lxY J< ( ((- $6  & gWNkeIQ^evc[^sQ|( N (lWxvy{|S~g"    010 c~gSR:N 4'Y{| 1 eO 6 $   )YQ(lx 7(lx )YQp 7(lp BBZ>( B2 eO ( t N(lx j(lx &; : ;3 Bgs(lWx   4 BgsN(lWx    #20 cRWVvg'`R"    g'`(lWx g'` N&^5uwN0ς0)YQp0 7(lp0j0JS0 g'`&^ck5uw)YQ070 g'`&^5uw~0V0|0g3 g$30 cNSO/f&TSTbegR  _(lWx:gSO NꁫTb _{1uߘirO~v(lWx0 V0r0ς0,02ukx0N0_N0N JS_(lWxNSOSTb FO|^?QgTb^ Nn0 ~*0|*0 ^_(lWx:gSOSꁫTbv(lWx0   q40^ˆ}v((lWxB &( $^ˆ}v((lWx_NwQ guir;m'`0P456$& N (lWxv͑tS'`(& $ 10N,irt'`( 20$N'`㉻yNI{5up$N'`'`(  1 (lWx:N$N'`5u( RM  # M' (2 aa㉻ye_SQN@bYsXvpH&$.  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N0ˆ}v(vuirf[R"  "66 H10RY7hˆ}v(RvY7h'` 1)0PS\O(u QNN@b gvv/fˆ}v(&Z Z%$ % 6 0o }\O(u \ }I{ 7 0cST OOo`vSSO SSOˆ}vI{ 8 0c6R~ހu0RS uVP[0;Oˆ}v 9 0%{Q\O(u0_;mRI{ebw͑\O(u0 20R YBgˆ}v(Rv YBg'` vPSRSYV }vq_T v^hsQzN'`0S'`I{ " dZ $ $ $  $($$)bN0ˆ}v(vR{|  "(  R{|SR cRP[b_r0RP[~b0n㉦^0uirR N cRP[b_rtrˆ}vT~~rˆ}v N cRP[~b{USˆ}vT~Tˆ}v  N cn㉦^nˆ}v0|ˆ}v0~ˆ}v0tˆ}v07ˆ}v0nˆ}v0lxˆ}vI{0 kZf f f k(V) cuirR* f;m'`ˆ}v(:Sb(Wu}TǏ z-NNR g;m'`vˆ}v(SvQMRSO0Y:o }ˆ}v(0Џˆ}v(0Obb2_ˆ}v(0kˆ}v(I{0 ^;m'`ˆ}v(ُ{|Wˆ}v([uirSOwObb/ec\O(u0Ylxˆ}v0҉ˆ}v0`Z$  $$  _  N0ˆ}v(v~b  "( N ˆ}v(vCQ }~b C0H0O0N S\ϑv S yrpN (WTyˆ}v(-Nvs^GW+Tϑ:N16% (u[.llSKm[ˆ}v(+Tϑ ˆ}v(+Tϑ(g) = ˆ}v(+T.lϑ6.25ff*$b@  V0(lWxvSf[  "( z(lWxamino acid, {Qaa) ˆ}v(vW,g~bUSMO /f+T(lWvx0 )Y6qˆ}v(1u20y(lWx~b0X$$9$$8 $N (lWxv~g  (  10_((( 20 -(lWxvgW  $gW1uNWVbSP[(WzzcRv N [y'` _wvIQf[;m'`zSO~g yzSOgW0 zSOgW (lW(W]:NL-W (lW(WS:ND-W/f=$f.N  6)Y6qpr-NvaaGW:NL-W $NWaavutR N T0 2^ v30 - (lWxwQeIQ'`du(lxY J< ( ((- $6  & gWNkeIQ^evc[^sQ|( N (lWxvy{|S~g"    010 c~gSR:N 4'Y{| 1 eO 6 $   )YQ(lx 7(lx )YQp 7(lp BBZ>( B2 eO ( t N(lx j(lx &; : ;3 Bgs(lWx   4 BgsN(lWx    #20 cRWVvg'`R"    g'`(lWx g'` N&^5uwN0ς0)YQp0 7(lp0j0JS0 g'`&^ck5uw)YQ070 g'`&^5uw~0V0|0g3 g$30 cNSO/f&TSTbegR  _(lWx:gSO NꁫTb _{1uߘirO~v(lWx0 V0r0ς0,02ukx0N0_N0N JS_(lWxNSOSTb FO|^?QgTb^ Nn0 ~*0|*0 ^_(lWx:gSOSꁫTbv(lWx0   q40^ˆ}v((lWxB &( $^ˆ}v((lWx_NwQ guir;m'`0P456$& N (lWxv͑tS'`(& $ 10N,irt'`( 20$N'`㉻yNI{5up$N'`'`(  1 (lWx:N$N'`5u( RM  # M' (2 aa㉻ye_SQN@bYsXvpH&$.  (!B3 I{5upisoelectric point, )Q:NpI 0"$f8 2ĉ[aaQ5uw:N:NevnmpHv:NaavI{5up.xfL *#5 I{5upv{&  $ 1u_Sw㉻y8^pe*    L 1udkSw ]w(WpIe [R+]= [R-] 4" tetS_'0  $NS[pe -lg [H+ ] 2 = - lg K1 - lg K2 5" -lg [H+ ] = pH - lg K1 = pK1 - lg K2 = pK2 4" 2 pH = pK1 + pK2 N pI:NI{5upevpH R pI =  pK1 +pK2 GlypI = 2 .4 + 9.6) = 5.97N                        #V1udkSw: I{5upvS_Naa$N'`yP[r`$NO WVpKevCO2S(uKmSlKmϑ {QSRS^vaaϑ0 c0ˆ}v(TYNSS^ FOupOe^]0Pr; 2 -& vQNSf[S^$ N2uS^ NN]xxvS^ NN]xWlDNFB vS^ _kx0lxovS^ C-zKm[ P56-57 NgIQvS^ N5 5 -SkxW-S(2-]xW2ux)vS^UU@ NN]xxvS^$XHN2 CH COOH +HNO2 R-CH-COOH + N2 R CH +H2O p 2    K *Nё^\yP[S^ ~B\g  2$V-Nv(lWx]^[teRP[ y(lWxkW(animo acid residue);6,$)Y6qX[(Wv;m'`$ (6 ;m'`(active peptide)X[(WNuirSOQ wQTyyrkuirf[Rv0 (O(;(;71mQ0ˆ}v(v~g" "( gWc(WzSO_gSO-NSNSP[bWV(WzzvST0 gacُNSNWV(WUS.eleSb_bv N TvzSO_gSO0 ˆ}v(vgacˆ}v(yr gvzz~g09n  M83N ˆ}v(vN~~g" $ $  ˆ}v(vN~~g/fN.ޏcaav~'`^R0 10T.v~g peptide)N*N(lWxv-WTSN(lWxv-(lW14l)T bvSTir0h'Z,Zff  !l 20vW,gޏce_  .( .(peptide bond):N1*N(lWxv-(lWTSN(lWxv-WKN14lTb_bvqQN. sSp.0 dk:Nˆ}v(-N(lWxޏcvW,ge_0(KZ;T  &z{ 1 .wQؚ^3z['` 2 wQS.'`( N1ulR 3 USCQHTs^bSTR'`'`( 4 C=ON NHS_cR0 AA6- yz 30/Qvޏc  Nkx.  i_1u-Nv^MO N$N*NJS(l x1"lޏc b dky(lxk W /fޏcQbv;N eh.0 R3z[zz~gKNN[u ir;mR0 :f ?:"Nkx.vޏc y{| gYy S'`BR$  @; O\ }"$0 -N\-~ހRlvNyˆ}v(o }0 RMNO@|0 ~g A21aaB30aa g$N[Nkx. 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